Pakistan-born terror suspect Tahawwur Hussain Rana, charged with helping to plan the November 26, 2008 terrorist attack in Mumbai that killed 166 people, has pleaded not guilty in federal court here.

Rana, 49, a Canadian citizen, wore an orange jumpsuit and entered the court room with both his hands and feet shackled. He spoke very briefly in the arraignment before Judge Arlander Keys on Monday, saying "Yes, your honour," when asked if he understood the charges against him.